Seeds remain in a dormant state, waiting for the right environmental conditions to germinate, thus increasing the probability of fragile seedling survival. The transition from a dry seed to a vegetative seedling is an irreversible process that requires a near-complete reprogramming of the plant transcriptome. While it is widely accepted that the activation of protein synthesis (translation) from seed-stored mRNAs is an essential step to trigger seed germination, much less was known about the re-initiation of the process of transcription.