NAC transcription factors (TFs) are involved in regulating complex signaling networks that respond to various abiotic stresses. However, it is unknown whether NAC TFs can negatively regulate fruit cold tolerance by regulating phospholipid degradation to produce PA. Thus, there is an urgent need to investigate the functions of cold-responsive NAC TFs in regulating phospholipid degradation to produce PA and to improve the understanding of plant responses to cold stress.